earlier cars had a pressurized expansion tank, it was part of the pressure side with the pressure cap on it. There is an overflow hose on it under the cap. The system is designed so that the fluid expands into the lines and tank, that way the rad is always full exposing maximum fluid level to cooling air.
The cap is a 2 way valve, it can suck in fluid from the external reservoir and pukes it there if the pressure climbs and the fludi expands over the rated pressure of the cap. If you look at the back, the outer rubber seal with the sprung cap is the pressure seal, that's what gives the cap it's rating. Inside it is an (vacuum) pressure valve that lets in fluid if the coolant in the pressurized part of the system contracts (cooling down). Without a reservoir (fed from bottom) you will push out hot coolant and when it contracts you will get a big air pocket in the upper part of the radiator. This is the way old downflow style rads were designed like used on fords and mopars well into the 70s. They only had a pressure cap w overflow hose, when running the rad is all full but check it cold and the fluid is well below the cap. These will spit out a good bit of fluid if you fill the rad on those to the brink when cold